Tremulus Lune debug – Please help! (voltages inside)

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I've built the Tremulus Lune (with the ramp switch) from the fuzzcentral layout at http://fuzzcentral.ssguitar.com/tremulus.php
I used the good old-fashioned cardboard circuit board method and have run into some problems...

Bypass works, and with the effect engaged the signal is coming through (and I can adjust the volume using the trimpot) but there is no effect. This is a minor improvement on 2 weeks ago when I first wired it up as there was no sound coming through at all with the stomp engaged... when I didn't get very far at all through the circuit with an audio probe, I put it away. I didn't change anything, but for some reason I now have signal. Odd, but I'm not complaining. ☺

Neither of the LED's light up at all. I used an LED/LDR combo as I couldn't get my hands on a Vactrol. I remember being very careful to orient the LEDs the correct way at the time... but I stranger things have happened I guess.

Anyway, some of the voltages I'm getting on my multimeter are as follows:

Battery: 8.88v

IC1: TL072

1. 3.05v
2. 3.05v
3. Jumps around a lot but eventually seems to settle on 5.09v
4. 0v
5. 2.84v
6. 3.77v
7. 8.03v
8. 8.69v

IC2: 4588

1. 4.44v
2. 4.43v
3. 4.22v
4. 0v
5. 4.21v
6. 4.42v
7. 4.41v
8. 8.83v

LDR: (+) 4.38v  (-) 4.38v
LED: (+) 3.05v  (-) 1.50v

(+) of 10uf cap (Triangle wave output – SMOOTH 3) eventually rises to 4.23v

Voltage on all electro caps is  higher on the (+) side.

After looking at a bunch of lune posts on the commonsound forum, I found a post that said voltages on pins 1, 2, 3 & 7 of the 4558 should be changing, (they don't) turning SPACING should change the pin 5 voltage, (it doesn't seem to) and SPEED should change pin 7 and the triangle wave output. (No luck there either.)

Is there anything in those voltages or my description that suggests something glaringly obvious that I've screwed up?

Any help is much appreciated!

EDIT: Corrected voltage for LED (-)